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Fermentation Atlas

Fish and seafood from Iceland

Hákarlhákarl

Greenland shark meat, which is toxic when fresh, is cut up, stacked in a draining box or a gravel pit and left for six to twelve weeks while bacteria break down its urea and TMAO into ammonia and trimethylamine. The pieces are then hung for weeks to months in an open drying shed and eaten in small cubes. Made in 3 steps over 2.5 months – 3.9 months. The longest step, stack and press to ferment (kæsing), takes 42 days – 2.8 months (Jensen S 2023) and runs at -3–20 °C (Jensen S 2023). Photobacterium (fresh meat) or Pseudoalteromonas (frozen meat), dominant in weeks 1–3 and linked to TMAO breakdown and Atopostipes, Pseudomonas and Tissierella do most of the fermenting.

Key numbers
Main step42 days – 2.8 monthsReported[matis2023]Stack and press to ferment (kæsing)
Temperature-3–20 °CMeasured[matis2023]Stack and press to ferment (kæsing)
AciditypH 6 → 8.9Measured[matis2023]Stack and press to ferment (kæsing)
Fully documentedFish and seafood2.5 months – 3.9 monthsMore fish and seafoodMore from Europe

Recipe

No fixed amounts recordedTraditional process with no added ingredients and no fixed batch size. The Matís trial used the meat of one 4–4.5 m female shark split into two equal halves.
Greenland shark meat, gutted and beheaded, cut into pieces · Back cuts and thinner belly (flap) cuts; the trial used 4–5 kg pieces[matis2023]
No salt and no starter · Nothing is added; bacteria come from the shark and its surroundings[matis2023][osimani2019]
Starter: spontaneous

Total time across the recorded steps: 2.5 months – 3.9 months

Start
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What changes: The gutted shark is cut into large pieces of meat

Step 1 of 3
Gut, behead and cut the shark
Time
Not documented
Temperature
Not documented
Salt
None given
Acidity
Not documented
Which microbes the steps name, step by step
Lactococcus main

In the Matís trial the shark was gutted and beheaded on board and kept on deck for two days before landing. Output: 4–5 kg back and flap cuts.

Flavour
Not documented
Texture
Not documented
Colour
Not documented

Microbes at work in this step

Not documented for this step.

Method

Measured measured in a studySet fixed by a specification or tested recipeReported one source reports itTypical a general value, not sourced for this fermentUnconfirmed not yet checked against a source

What makes this one different

Hákarl is fermented with no salt at all and turns alkaline rather than sour; the main change is bacteria turning the shark's urea and TMAO into ammonia and trimethylamine. Unlike surströmming it is finished by air-drying, not canning.

Related ferments

Variants

  • Glerhákarl"Glassy shark": chewy, reddish belly meat.
  • Skyrhákarl"Skyr shark": soft, white body meat.

Safety

Fresh Greenland shark meat is considered toxic if eaten in quantity because it holds a lot of urea and TMAO; fermentation turns these into ammonia, trimethylamine and dimethylamine, so it is both preservation and detoxification. The alkaline pH of 8–9 holds back spoilage microbes, but finished hákarl has water activity near 0.96, enough for an active microbial community. Shark frozen before fermenting started with more TMAO but ended with less TMA and DMA.

Hazards the sources name: Listeria (minor DNA sequences, under 1 %, in ready-to-eat samples); Shiga toxin-producing Escherichia coli (tested for; not found).

History and sources

Every number is labelled with how it is known. How this atlas is made

The oldest record of processed shark in Iceland dates from 1374. Hákarl was long a valued source of protein and energy, and today it is eaten mainly at the midwinter Þorri feasts.

  1. [matis2023] Jensen S., Ólason S.P., Skírnisdóttir S., Stefánsson G., Dargentolle C., Marteinsson V.T. 2023 (Matís), Unlocking the microbial diversity and the chemical changes throughout the fermentation process of hákarl, Greenland shark, Heliyon 9:e22127paper
  2. [osimani2019] Osimani A. et al. 2019, Unveiling hákarl, a study of the microbiota of the traditional Icelandic fermented fish, Food Microbiologypaper
  3. [cardinali2026] Cardinali F. et al. 2026, Novel insights into hákarl, a deep dive into the microbiological and physico-chemical features of Iceland's traditional fermented shark, International Journal of Food Microbiology 111629paper
  4. [matis-web] Matís, Unlocking the microbial diversity and chemical changes during fermentation of hákarl (summary page)web